This advanced course offers students the opportunity to study the unique biodiversity of the Galapagos Islands, where the delicate balance between geological, biological, climatologic and anthropogenic processes in controlling the process of evolution is on display. The Galapagos are a unique locale where the fragile nature of the environment, the processes that shape it, and the unique nature of its inhabitants are readily observed. Additionally, the historic significance of this locale in the development of one of the most fundamental of all scientific theories will be explored fully.